1. ISEM, CNRS, Univ. of Montpellier, IRD, EPHE Montpellier France
2. School of Animal, Plant and Environmental Science, Univ. of the Witwatersrand Wits South Africa
3. Museum of Comparative Zoology, Dept of Organismic and Evolutionary Biology, Harvard Univ Cambridge MA USA
4. Dept of Mammalogy, National Museum, and Centre for Environmental Management, Univ. of the Free State Bloemfontein South Africa
5. Dept of Economic Development, Tourism and Environmental Affairs, Biodiversity Research Bloemfontein South Africa
6. Centre de Coopération Internationale en Recherche Agronomique pour le Développement (CIRAD), UMR, CMAEE, C2B Montpellier France