1. Pearl River Fisheries Research Institute Chinese Academy of Fishery Science Guangzhou China
2. Key Laboratory of Aquatic Animal Immune Technology of Guangdong Province Guangzhou China
3. Guangzhou Scientific Observing and Experimental Station of National Fisheries Resources and Environment Guangzhou China
4. Institut de Recherche pour le Développement, UMR 226 ISEM (UM‐CNRS‐IRD) Montpellier France
5. Hainan Academy of Ocean and Fisheries Sciences Haikou China
6. College of Fisheries Henan Normal University Xinxiang China