1. School of Plant Biology; The University of Western Australia; Perth Australia
2. The UWA Institute of Agriculture; The University of Western Australia; Perth Australia
3. The Southern Grassland Ecosystem Research Station; South China Botanical Garden, Chinese Academy of Sciences; Guangzhou 510650 China
4. School of Earth and Environment; The University of Western Australia; Perth Australia
5. Department of Environmental Science and Technology; School of Applied Sciences, Cranfield University; Cranfield UK