1. Lewis B. and Dorothy Cullman Program for Molecular Systematic Studies; The New York Botanical Garden; Bronx New York 10458 USA
2. Department of Botany; Academy of Natural Sciences; Philadelphia Pennsylvania 19103 USA
3. Department of Plant Science and Landscape Architecture; University of Maryland; College Park Maryland 20742 USA
4. Department of Biodiversity, Earth, and Environmental Sciences; Drexel University; Philadelphia Pennsylvania 19104 USA