1. Systematic Biology, Department of Organismal Biology Uppsala University Uppsala Sweden
2. Institut de Biochimie et de Génétique Cellulaire, UMR 5095 CNRS Université de Bordeaux Bordeaux CEDEX France
3. Department of Biology, Realfagbygget Norwegian University of Science and Technology Trondheim Norway
4. Division of Population Genetics, Department of Zoology Stockholm University Stockholm Sweden