1. Department of Biological Sciences; Tokyo Metropolitan University; Minamiosawa 1-1 Hachioji 192-0397 Japan
2. Department of Population Genetics; National Institute of Genetics; 1111 Yata Mishima 411-8540 Japan
3. Department of Genetics; SOKENDAI (The Graduate University for Advanced Studies); 1111 Yata Mishima 411-8540 Japan
4. Research Center for Genomics and Bioinformatics; Tokyo Metropolitan University; Minamiosawa 1-1 Hachioji 192-0397 Japan