1. School of Biological, Earth and Environmental Science; University of New South Wales; Sydney NSW 2052 Australia
2. Department of Sustainability, Environment, Water, Population and Communities; Australian Antarctic Division; Kingston TAS 7050 Australia
3. School of Biological Sciences; University of Sydney; Sydney NSW 2006 Australia
4. Antarctic Climate and Ecosystems Cooperative Research Centre; University of Tasmania; Hobart TAS 7001 Australia