1. Fish Ecology and Conservation Physiology Laboratory, Department of Biology and Institute of Environmental and Interdisciplinary Science Carleton University Ottawa Ontario Canada
2. Department of Biology Trent University Peterborough Ontario Canada
3. Ohio Department of Natural Resources, Division of Wildlife Sandusky Fisheries Research Station Sandusky Ohio USA
4. Great Lakes Acoustic Telemetry Observation System, Department of Fisheries and Wildlife Michigan State University East Lansing Michigan USA