1. College of Life Sciences; Sichuan University; Chengdu 610064 China
2. Department of Plant and Microbial Biology; University of California at Berkeley; Berkeley CA 73072 USA
3. NJU-NJFU Joint Institute for Plant Molecular Biology; College of Life Sciences; Nanjing University; Nanjing 210093 China
4. Ministry of Education Key Laboratory of Cell Activities and Stress Adaptations; School of Life Sciences; Lanzhou University; Lanzhou 730000 China