1. College of Horticulture Science and Engineering Shandong Agricultural University Tai’an China
2. Collaborative Innovation Center of Fruit & Vegetable Quality and Efficient Production Shandong China
3. Key Laboratory of Biology and Genetic Improvement of Horticultural Crops in Huanghuai Region Ministry of Agriculture and Rural Affairs Tai’an China
4. State Key Laboratory of Crop Biology Tai'an China