1. Division of Infectious Animal Diseases National Institute of Animal Health Ibaraki Japan
2. Laboratory of Veterinary Hygiene, Faculty of Veterinary Medicine, Graduate School of Infectious Diseases, and One Health Research Center Hokkaido University Hokkaido Japan
3. Institute of Natural and Environmental Sciences University of Hyogo/Wildlife Management Research Center Hyogo Japan
4. Faculty of Medicine University of Miyazaki Miyazaki Japan