1. Biodiversity and Conservation Science Department of Biodiversity Conservation and Attractions, Locked Bag 104, Bentley Delivery Centre Bentley WA 6983 Australia
2. School of Biological Sciences, The University of Western Australia 35 Stirling Highway, Crawley WA 6009 Australia
3. Kings Park Science, Department of Biodiversity Conservation and Attractions Kattidj Close, Kings Park, WA 6005 Australia
4. Centre of Excellence in Natural Resource Management, School of Agriculture and the Environment, The University of Western Australia 35 Stirling Terrace, Albany WA 6330 Australia