1. Department of Benthic Ecology; GEOMAR Helmholtz-Zentrum für Ozeanforschung Kiel; 24105 Kiel Germany
2. College of Marine Life Sciences; Ocean University of China; Qingdao 266003 China
3. Xunshan Group Co., Ltd; Rongcheng 264316 China
4. Akkeshi Marine Station; Field Science Center for Northern Biosphere; Hokkaido University; Akkeshi Hokkaido 088-1113 Japan