1. Department of Biology; Boston University; 5 Cummington Mall Boston MA 02215 USA
2. Cary Institute of Ecosystem Studies; Box AB; Millbrook NY 12545 USA
3. Bates College; 7 Andrews Road Lewiston ME 04240 USA
4. Department of Integrative Biology; University of California; Berkeley CA 94720 USA
5. Department of Environmental Science; Policy & Management; University of California; Berkeley CA 94720 USA