1. Ecology and Biodiversity; Institute of Environmental Biology; Utrecht University; Padualaan 8 3584 CH Utrecht The Netherlands
2. School of Biological Sciences; Centre for Biodiversity and Conservation Science; The University of Queensland; St Lucia Qld 4072 Australia
3. Evolutionary Demography Laboratory; Max Planck Institute for Demographic Research; DE-18057 Rostock Germany
4. Department of Zoology; School of Natural Sciences; Trinity College Dublin; Dublin 2 Ireland
5. Forest Ecology and Management; Wageningen Universiteit, Lumen; Droevendaalsesteeg 3a Postbus 47 6700 AA Wageningen The Netherlands