1. Coastal Ecosystems Research Station of Yangtze River Estuary; Ministry of Education Key Laboratory for Biodiversity Science and Ecological Engineering; Institute of Biodiversity Science; Fudan University; #220 Handan Road Shanghai 200433 China
2. Department of Plant Pathology and Ecology; The Connecticut Agricultural Experiment Station; New Haven CT 06504 USA
3. School of Life Sciences; University of KwaZulu-Natal; Private Bag X 01 Scottsville 3209 South Africa