1. Department of Ecology and Evolution; University of Lausanne; Biophore Building 1015 Lausanne Switzerland
2. Department of Ecology and Evolutionary Biology; University of California; 321 Steinhaus Hall Irvine CA 92697 USA
3. Department of Biology; Unit of Ecology & Evolution; University of Fribourg; Ch. du Musée 10 CH-1700 Fribourg Switzerland