1. Forestry and Forest Products Research Institute Tsukuba Japan
2. CAS Key Laboratory of Forest Ecology and Management Institute of Applied Ecology Chinese Academy of Sciences Shenyang China
3. Organismal and Evolutionary Biology Viikki Plant Science Centre (ViPS) University of Helsinki Helsinki Finland
4. UNIROUEN URA/EA1293 IRSTEA FR Scale CNRS 3730 Normandie Université Rouen France
5. Graduate School of Life Sciences Tohoku University Sendai Japan
6. Department of Botany National Museum of Nature and Science Tsukuba Ibaraki Japan