1. Ecology & Biodiversity Group; Institute of Environmental Biology; Utrecht University; Padualaan 8 3584 CH Utrecht The Netherlands
2. Dutch Centre for Avian Migration and Demography; Netherlands Institute of Ecology (NIOO-KNAW); Droevendaalsesteeg 10 6708 PB Wageningen The Netherlands
3. Department of Animal Ecology; Netherlands Institute of Ecology (NIOO-KNAW); Droevendaalsesteeg 10 6708 PB Wageningen The Netherlands
4. Theoretical and Computational Ecology; IBED; University of Amsterdam; Amsterdam The Netherlands