1. State Key Laboratory of Water Environment Simulation School of Environment Beijing Normal University Beijing China
2. Conservation Ecology Group Groningen Institute for Evolutionary Life Sciences University of Groningen Groningen the Netherlands
3. Yellow River Estuary Wetland Ecosystem Observation and Research StationMinistry of Education Shandong China
4. Department of Estuarine and Delta Systems Royal Netherlands Institute for Sea Research and Utrecht University Yerseke the Netherlands