1. State Key Laboratory of Forest and Soil Ecology; Institute of Applied Ecology; Chinese Academy of Sciences; Shenyang Liaoning 110164 China
2. School of Biology; Georgia Institute of Technology; Atlanta GA 30332 USA
3. Forest dynamics; Swiss Federal Research Institute WSL; CH-8903 Birmensdorf Switzerland
4. Nature Conservation and Plant Ecology Group; Wageningen University; PO Box 47 6700 AA Wageningen The Netherlands
5. State Key Laboratory of Vegetation and Environmental Change; Institute of Botany; Chinese Academy of Sciences; Beijing 100093 China
6. Key Laboratory of Plant Stress Biology; College of Life Sciences; Henan University; Kaifeng Henan 475004 China
7. Department of Biology and Graduate Degree Program in Ecology; Colorado State University; Fort Collins CO 80523 USA