1. State Key Laboratory of Vegetation and Environmental Change; Institute of Botany; the Chinese Academy of Sciences; Beijing China
2. Key Laboratory of Ecosystem Network Observation and Modeling; Synthesis Research Center of Chinese Ecosystem Research Network; Institute of Geographic Sciences and Natural Resources Research; Chinese Academy of Sciences; Beijing 100101 China
3. Environmental Studies; University of California; Santa Cruz CA 95064 USA
4. State Key Laboratory of Forest and Soil Ecology; Institute of Applied Ecology; Chinese Academy of Sciences; Shenyang China