1. School of Natural Resources and Environment; University of Michigan; Ann Arbor MI 48109-1041 USA
2. Section of Earth and Environmental Sciences; Institut F.-A. Forel; University of Geneva; 10 Route de Suisse; CP 416 1290 Versoix Switzerland
3. Department of Ecology, Evolution, and Marine Biology; University of California; Santa Barbara CA 93106-9620 USA