1. State Key Laboratory of Vegetation and Environmental Change Institute of Botany, Chinese Academy of Sciences Beijing China
2. University of the Chinese Academy of Sciences Beijing China
3. Northeast Institute of Geography and Agroecology Chinese Academy of Sciences Changchun China
4. Department of Biology University of Kentucky Lexington Kentucky USA
5. Department of Plant and Soil Sciences University of Kentucky Lexington Kentucky USA
6. School of Life Sciences Hebei University Baoding China
7. School of Biological Sciences Georgia Institute of Technology Atlanta Georgia USA