1. Long-term Ecology Laboratory, Biodiversity Institute, Department of Zoology; University of Oxford; Oxford; OX1 3PS; UK
2. Island Ecology and Biogeography Group, Instituto Universitario de Enfermedades Tropicales y Salud Pública de Canarias (IUETSPC); University of La Laguna; La Laguna; 38206; Canary Islands; Spain