1. Graduate School of Environmental Science Hokkaido University North 10 West 5, Kita‐Ku Sapporo 060‐0810 Japan
2. Institute for Marine and Antarctic Studies University of Tasmania 20 Castray Esplanade, Battery Point Hobart TAS 7004 Australia
3. South Australian Research and Development Institute 2b Hartley Grove Urrbrae SA 5064 Australia
4. Faculty of Environmental Earth Science Hokkaido University North 10 West 5, Kita‐Ku Sapporo 060‐0810 Japan