1. Department of Organismic and Evolutionary Biology; Harvard University Herbaria; Harvard University; Cambridge MA 02138 USA
2. Biology Department; Boston University; Boston MA 02215 USA
3. Pringle Herbarium; Plant Biology Department; University of Vermont; Torrey Hall, 27 Colchester Ave Burlington VT 05405 USA
4. Brown University Herbarium; Department of Ecology and Evolutionary Biology; Brown University; 34 Olive Street, Box G-B225 Providence RI 02912 USA
5. Biology Department; University of Massachusetts; 611 North Pleasant Street Amherst MA 01003 USA
6. Division of Botany; Peabody Museum of Natural History; Yale University; New Haven CT 06511 USA
7. Harvard Forest; Harvard University; 324 North Main Street Petersham MA 01366 USA
8. Tropical Forests & People Research Centre; University of the Sunshine Coast; Maroochydore Qld 4558 Australia