1. Nottingham Digestive Diseases Biomedical Research Centre, School of Medicine University of Nottingham Nottingham UK
2. NIHR Nottingham Biomedical Research Centre Nottingham University Hospitals NHS Trust and the University of Nottingham Nottingham UK
3. Public Health England, Seaton House Nottingham UK
4. Population and Lifespan Sciences, School of Medicine University of Nottingham Nottingham UK