1. School of Environment and Natural Resources, The Ohio State University; 2021 Coffey Road Columbus OH 43210 USA
2. North Appalachian Experimental Watershed, US Department of Agriculture, Agricultural Research Service; PO Box 488 Coshocton OH 43812 USA
3. School of Agriculture, Food and Environment, The Royal Agricultural University; Cirencester Gloucestershire GL7 6JS UK
4. Sustainable Soils and Grassland Systems Department; Rothamsted Research; North Wyke, Okehampton Devon EX20 2SB UK