1. State Key Laboratory of Plant Genomics, Center for Agricultural Resources Research, Institute of Genetics and Developmental Biology Chinese Academy of Sciences Shijiazhuang Hebei 050021 China
2. School of Agriculture and Biology Shanghai Jiao Tong University 800 Dongchuan Road, Minhang District Shanghai 200240 China
3. Key Laboratory of Molecular and Cellular Biology of Ministry of Education, Hebei Collaboration Innovation Center for Cell Signaling, Hebei Key Laboratory of Molecular and Cellular Biology, College of Life Sciences Hebei Normal University Shijiazhuang 050024 China
4. University of Chinese Academy of Sciences Beijing 100049 China