1. Department of Forestry and Environmental Resources, Program in Fisheries, Wildlife and Conservation Biology North Carolina State University Raleigh NC 27695‐7617 U.S.A.
2. Wildlife Biology Program University of Montana Missoula MT 59812 U.S.A.
3. Bureau of Land Management Missoula Field Office Missoula MT 59804 U.S.A.
4. Wildlife Biology Program and Office of the Vice President for Research and Creative Scholarship University of Montana Missoula MT 59812 U.S.A.