1. Université Montpellier 2; Montpellier France
2. CNRS-UMR5554 Institut des Sciences de l'Evolution; Station Méditerranéenne de l'Environnement Littoral; Sète France
3. Department of Ecology and Evolution; University of Lausanne; Lausanne Switzerland
4. Université Lille Nord de France; USTL; GEPV; CNRS; FRE 3268; Villeneuve d'Ascq France
5. Department of Ecology and Evolutionary Biology; University of California; Santa Cruz CA USA