1. School of Life and Environmental Sciences Deakin University Geelong Victoria Australia
2. School of Life and Environmental Sciences Deakin University Burwood Victoria Australia
3. CREEC/CANECEV (CREES), MIVEGEC IRD 224–CNRS 5290–Université de Montpellier Montpellier France
4. Evolution & Ecology Research Centre, School of Biological, Earth and Environmental Sciences University of New South Wales Sydney New South Wales Australia