1. The Centre for Environment, Fisheries & Aquaculture Science; Lowestoft Laboratory; Lowestoft, Suffolk NR33 0HT U.K.
2. Game & Wildlife Conservation Trust, Salmon and Trout Research Centre; The River Laboratory; East Stoke, Wareham, Dorset BH20 6BB U.K.
3. Queen Mary University of London, School of Biological Sciences; Mile End Road London, E1 4NS U.K.