1. School of Biological Sciences; Flinders University; Adelaide South Australia 5001 Australia
2. South Australian Research and Development Institute (SARDI); Aquatic Sciences Centre; Adelaide South Australia 5024 Australia
3. Centre National de la Recherche Scientifique, Laboratoire d'Océanologie et de Géosciences; UMR 8187, Université des Sciences et Technologies de Lille; Station Marine Wimereux 62930 France