1. Division of Biomedical and Life Sciences, Faculty of Health and Medicine Lancaster University Lancaster UK
2. MRC Human Genetics Unit, Institute of Genetics and Cancer, Western General Hospital University of Edinburgh Edinburgh UK
3. Centre for Skin Sciences, Faculty of Life Sciences University of Bradford Bradford UK
4. Cancer Research UK, Beatson Institute, and Institute of Cancer Sciences University of Glasgow Glasgow UK
5. Roslin Institute University of Edinburgh Roslin UK