1. Department of Ecology, Evolutionary and Organismal Biology; Iowa State University; Ames IA 50011 USA
2. Department of Ecology and Evolutionary Biology; University of Arizona; Tucson AZ 85721 USA
3. BioCircuits Institute; University of California, San Diego; La Jolla CA 92093-0328 USA
4. Department of Biological Science; University of Pittsburgh; Pittsburgh PA 15260 USA
5. School of Biological Sciences; Royal Holloway University of London; Surrey TW20 0EX UK
6. Department of Environmental Science and Policy; University of California; Davis CA 95616 USA