1. Centre for Ecology, Evolution and Conservation School of Biological Sciences University of East AngliaNorwich Research Park Norwich UK
2. Groningen Institute for Evolutionary Life Sciences Faculty of Science and Engineering University of Groningen Groningen The Netherlands
3. Department of Animal and Plant Sciences University of Sheffield Sheffield UK
4. School of Biology Faculty of Biological Sciences University of Leeds Leeds UK
5. Nature Seychelles Mahe Republic of Seychelles