1. State Key Laboratory for Conservation and Utilization of Subtropical Agro-bioresources; College of Life Sciences, South China Agricultural University; Guangzhou 510642 China
2. Shenzhen Key Laboratory of Southern Subtropical Plant Diversity, Fairylake Botanical Garden; Shenzhen and Chinese Academy of Sciences; Shenzhen 518004 Guangdong China
3. Marine Environmental Monitor Center of Shanwei; State Oceanic Administration; Shanwei 516600 Guangdong China
4. State Key Laboratory of Systematic and Evolutionary Botany; Institute of Botany, Chinese Academy of Sciences; Beijing 100093 China
5. Hengshan National Nature Reserve Administration; Hengshan 421900 Hunan China