1. Department of Marine Ecosystems Dynamics; Atmosphere and Ocean Research Institute; The University of Tokyo; 5-1-5 Kashiwanoha Kashiwa Chiba 277-8564 Japan
2. SNSB-Bavarian State Collection of Zoology; Münchhausenstr. 21 81247 Munich Germany
3. Department Biology II; Ludwig-Maximilians-Universität; Großhaderner Str. 2 82152 Planegg-Martinsried Germany