1. Department of Biology St. Mary's College of Maryland St. Mary's City MD 20653 USA
2. Division of Biological Sciences University of Montana Missoula MT 59812 USA
3. School of Forestry Northern Arizona University Flagstaff AZ 86011 USA
4. University and Jepson Herbaria, and Department of Integrative Biology University of California Berkeley Berkeley CA 94720‐2465 USA
5. School of Life Sciences University of Nevada Las Vegas Las Vegas NV 89154 USA