1. Shandong Provincial Key Laboratory of Energy Genetics and CAS Key Laboratory of Biofuels Qingdao Institute of Bioenergy and Bioprocess Technology Chinese Academy of Sciences Qingdao 266101 China
2. University of Chinese Academy of Sciences Beijing 100049 China
3. CAS Key Laboratory of Tibetan Medicine Research Northwest Institute of Plateau Biology Chinese Academy of Sciences Xining 810008 China
4. Sichuan Academy of Grassland Science Chengdu 611731 China