1. Department of Evolutionary Biology and Environmental Studies University of Zurich Zurich8057 Switzerland
2. Department Biology University of Cadiz Puerto Real 11510 Spain
3. Department of Zoology Oxford University New Radcliffe House Radcliffe Observatory Quarter Woodstock Rd OxfordOX2 6GGUK
4. Department of Animal & Plant Sciences University of Sheffield Alfred Denny Building, Western Bank SheffieldS10 2TN UK
5. Centre for Biodiversity and Conservation Science University of Queensland St Lucia4071 Qld. Australia
6. Evolutionary Demography Laboratory Max Plank Institute for Demographic Research Rostock18057 Germany