1. Department of Biological Sciences California State University Sacramento CA USA
2. Smithsonian Tropical Research Institute Apartado 0843‐03092 Ancon Panama
3. Department of Botany University of Hawaii at Manoa Honolulu HI USA
4. Department of Environmental Engineering Sciences Engineering School of Sustainable Infrastructure and Environment University of Florida Gainesville FL32611 USA
5. Smithsonian Environmental Research Center Edgewater MD USA