1. Institut des Sciences de l'Evolution de Montpellier (Université de Montpellier | CNRS | IRD | EPHE); Montpellier France
2. Department of Biological Sciences; University of Alberta; Edmonton Canada
3. INRA, UMR 1062 Centre de Biologie pour la Gestion des Populations (INRA, IRD, CIRAD, Montpellier SupAgro); Montferrier-sur-Lez France
4. Department of Plant and Environmental Protection Sciences; University of Hawai'i at Mānoa; Honolulu HI U.S.A.