1. Department of Physical Geography; Utrecht University; Utrecht The Netherlands
2. Faculty of Veterinary Medicine; Utrecht University; Utrecht The Netherlands
3. Ecology Evolution and Genomics of Infectious Disease Research Group; Institute of Integrative Biology; The University of Liverpool; Liverpool UK
4. Evolutionary Ecology Group; Department of Biology; University of Antwerp; Antwerp Belgium
5. Stavropol Plague Control Research Institute; Stavropol Russian Federation