1. College of Biological Sciences and Technology Beijing Forestry University Beijing 100083 China
2. National Engineering Laboratory for Tree Breeding Beijing Forestry University Beijing 100083 China
3. Agricultural and Biological Engineering Department University of Florida Gainesville FL 32611 USA
4. College of Life Science Linyi University Linyi 276005 China
5. Key Laboratory of Tree Breeding and Cultivation State Forestry Administration Research Institute of Forestry Chinese Academy of Forestry Beijing 100091 China