1. Department of Biology, Faculty of Science Kyushu University Fukuoka Japan
2. Department of Botany National Museum of Nature and Science Tsukuba Ibaraki Japan
3. Graduate School of Life Sciences Tohoku University Sendai Japan
4. Iriomote Station, Tropical Biosphere Research Center University of the Ryukyus Okinawa Japan
5. Biodiversity Division National Institute for Environmental Studies Tsukuba Ibaraki Japan
6. College of Arts and Sciences J. F. Oberlin University Machida Tokyo Japan