1. Joint International Research Laboratory of Metabolic & Developmental Sciences, State Key Laboratory of Hybrid Rice, School of Life Sciences and Biotechnology Shanghai Jiao Tong University Shanghai 20040 China
2. Department of Energy Plant Research Laboratory and Plant Biology Department Michigan State University East Lansing MI 48824 USA
3. State Key Laboratory of Hybrid Rice, Hunan Hybrid Rice Research Center Hunan Academy of Agricultural Sciences Changsha 410125 China