1. Laboratory of Animal Breeding and Genetics; Graduate School of Biosphere Science; Hiroshima University; Higashi-Hiroshima Hiroshima Japan
2. Japanese Avian Bioresource Project Research Center; Hiroshima University; Higashi-Hiroshima Hiroshima Japan
3. Institute of Laboratory Animals; Graduate School of Medicine; Kyoto University; Sakyou-ku Kyoto Japan